The Movius Line represents the crossing of a demographic threshold

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When examining the dispersal of Pleistocene hominins, one of the more fascinating debates concern the patterns of biological and technological evolution in East Asia and other regions of the Old World. One suggestion emerging from palaeoanthropological research places a demarcation between these two regions in the form of a geographical division known as the Movius Line. Specifically, the suggestions that initially led to the Movius Line were based on observations of differing technological patterns, namely: the lack of Acheulean handaxes and the Levallois core traditions in East Asia.

Since Hallam L. Movius’ initial proposal, the recent discovery of handaxes within East Asia have led to suggestions that the Movius Line is in fact obsolete. Suggesting this may not in fact be the case is a recent paper by Stephen Lycett & Christopher Norton, which highlights three central points coming from a growing body of research: 1) “several morphometric analyses have identified statistically significant differences between the attributes of specific biface assemblages from east and west of the Movius Line”; 2) “The number of sites from which handaxes have been recovered in East Asia tend to be geographically sparse compared with many regions west of the Movius Line”;  3) “‘handaxe’  specimens  tend only  to comprise a  small percentage of the total number of artefacts recovered, a situation that  contrasts  with  many  classic  Acheulean  sites  in  western portions of the Old World, where bifacial handaxes may dominate assemblages in large numbers”.

In light of these developments, the current paper combines cultural transmission theory and demography to produce a “generalised model for Palaeolithic technological evolution during the Pleistocene”. As it is generally assumed, cultural transmission underlies technological traditions, which may be taken as “a particular behaviour (e.g., tool manufacture and use) that is repeated over generations, and is learned and passed on between individuals via a process of social interaction”. Furthermore, we now also know such forms of transmission can be modelled in an analogous manner to that of genetic transmission. In fact, the parallels between genetic and cultural transmission extend to the point where “factors known to structure patterns of genetic variation and transmission (e.g., drift, selection, dispersal and demography) must also be taken into account when examining patterns of cultural variation across space and time”.

Like in population genetics, Lycett & Norten refer to effective population size (Ne), except instead of meaning the number of individuals actively involved in passing on genetic material, they use the term to mean “the number of skilled practitioners of a given craft tradition involved in passing on those skills to subsequent generations via social transmission”. As such, small effective populations rely more on stochastic factors, such as drift, in shaping which cultural variants will be passed on to future generations. By contrast, large populations are less likely to be impacted upon by stochastic sampling effects – allowing for a greater number of innovations to diffuse throughout the population. This is supported via mathematical modelling, which shows:

[...] that a decrease in effective population size (Ne) may lead to a loss of pre-existing socially transmitted cultural elements… [therefore] the greater the number of models, the more choice is available for selecting the best (i.e. most skilled) models from which to copy. That is, in larger populations, cumulative cultural learning is possible because the effect of having a larger number of models from which to pick the most skilled, exceeds the losses resulting from imperfect copying of that skill. Hence, the chance of copying the most skilled elements of a given practice correlates directly with the number of models from which to copy.

An example of these demographic conditions in action comes from the Tasmanian Islanders. When Tasmania became isolated from Australia approximately 10-12 thousand years ago, the newly established islanders appear to have lost and/or never developed the ability to manufacture a range of technologies, including: fishing spears, cold-weather clothing and boomerangs. Described as a ‘cultural founder effect’, Lycett & Norton use these observations to implement the following three conditions in constructing their model: population size, density and social interconnectedness:

Social interconnectedness reflects the likelihood of encountering a given craft skill and the regularity of such encounters. Social interconnectedness is thus somewhat proportional to the parameters of effective population size (i.e. number of skilled craft practitioners) and population density (i.e. probability of encounter due to degree of aggregation).

By varying these population conditions, Lycett and Norton use their model to show how different stages of lithic technologies may be sustained (see figure below, taken from Lycett & Norton(2010)). Crucially, demographic levels may decrease and lead to a situation where sustaining already-created technological innovations may not be tenable for the long-term. In providing a null model, they remain neutral as to the effects of cognitive and biomechanical evolution on the emergence and disappearance of technological patterns. In fact, the demographic processes described (larger populations and greater density) mirror those conditions under which human biological evolution may have accelerated. Meaning, human population structures are advantageous for the spread of novel technologies, cultural variants and adaptive genes. This may have implications for why we are seeing growing evidence of gene-culture coevolution in modern human populations.

Although it’s prohibitively difficult to estimate the demographic parameters of ancient hominin populations, it is fairly certain that demographic levels will have been relatively higher within Africa than in other parts of the world during the Early to Middle Pleistocene. When coupled with evidence showing the earliest First Appearance Dates (FADs) for Mode 1, Mode 2 and Mode 3 technologies in Africa, the demographic model presented is certainly congruent with the assertion that there is a definitive link between the spread of technological innovations and sustained population growth. Thus, if hominin populations were temporally and spatially discontinuous, then they simply lacked the necessary population conditions to maintain a certain degree of technological sophistication. By extension, explaining the relative absence of bifacial implements in East Asia is perhaps a demographic problem.

As mentioned earlier, we do see some evidence for Acheulean technology in East Asia. But this is not a refutation of the model. Rather, the discovery of these technologies perfectly fits within the model, especially as it is not at any significant level when compared to those found west of the Movius Line. That is, even if Acheulean technologies were imported to East Asia, the demographic conditions would be unfavourable in maintaining them. In this case, we would expect to see Acheulean technology at relatively low densities in East Asia. As the authors note:

One further observation that is not often noted is that the artefact density of most of the Early Palaeolithic sites in East Asia is also usually very low. For instance, in Fangniushan and Chenshan, two Middle Pleistocene open-air sites in central-east China, the artefact densities are less than one per m3

Of course, artefact density may be influenced by a whole host of factors. Raw material availability is one example. More broadly, the markedly different technological patterns observed may not be causally related to demography at all. We could find evidence for sharp cognitive differences between hominin populations east and west of the Movius Line. However, the  authors predict that these explanations will become increasingly problematic if “site densities and the chronological distribution of sites in East Asia [...] continue to differ from those in the west”.

But this is what’s great about null models: they are testable. In this case, the model predicts that:

[...] evidence for demographic levels in East Asia will be found to be significantly different from those in many parts of western Eurasia and Africa during the Early and Middle Pleistocene. Here, we have hinted at some of the currently available evidence that suggests this may have been the case. What is now urgently  needed  are  more  sophisticated  means  than  we  have provided here of assessing Pleistocene demographic parameters in the key regions east and west of the Movius Line.

So it’s still very much an open question as to why we find differing technological patterns east and west of the Movius Line. But demography certainly seems like a good candidate.

Citation: Lycett, S.J & Norton, C.J. A demographic model for Palaeolithic technological evolution: The case of East Asia and the Movius Line. Quaternary International, 211 (1-2), 55-65 DOI: 10.1016/j.quaint.2008.12.001.

8 Comments

  1. We covered this in my graduate seminar this week. so maybe I’ll write something about it. I think in the end it falls apart on the differential equation level: the rate of introduction of bifaces is fairly high (that’s why we’ve found them in China and Korea, now), and they persisted at least long enough for us to have found them. So if they were locally useful they had the chance to spread, but they didn’t. That argues for an ecological reason, that they just weren’t useful in the East.

    Oh, and there are those Flores hominids, making tools for hundreds of thousands of years in a tiny population, and Java periodically cut off from SE Asia. Toolmaking seems like it wasn’t very easy to lose.

  2. Prof. Hawks, what do you think about the contrast between Levallois (Shuidonggou) and non-Levallois (Xiachuan) cultures/blade technologies existing close by. Is this a case of control origin but lost of transmission, cultural spread of blade technology but not Levallois, or parallel adaptation, in your opinion?

  3. Sorry, in the comment above I meant “common origin” but typed “control origin”.

  4. Good question, I don’t think anybody has a very good idea. It’s probably a mistake to think of Levallois (or blades) as cases of unique invention. They are recurrently popping up, even in Lower Paleolithic contexts. If you thought about tools with a genetic analogy, the appearance of these techniques is not “mutation-limited” — that is, populations didn’t have to wait around for somebody to invent them.

    The question is, if there’s some optimum, why don’t people make the same tools everywhere? And you’ve got four possible answers:

    (1) Different optima; different ecological contexts or raw material availability require different tools.

    (2) Transmission is too hard to keep going in these populations.

    (3) Path-dependence: slightly different starting points may lead to very different technical developments.

    (4) Relative neutrality: It just doesn’t matter very much which tools you use.

    Path-dependence is probably the most common explanation for why Levallois is rare in China — the idea being that you don’t develop prepared cores easily if you aren’t starting from a large cutting tool tradition. What isn’t easily explained is why sites after 50,000 years ago should be susceptible to this same path-dependence — particularly since they should derive some or most of their ancestry from Africa/West Asia.

    Personally, I keep coming back to ecology. For two traditions relatively late in the record and possibly synchronous (although I hesitate to assume this until a better chronology emerges), I tend to go with Binford — they’re probably doing different things.

  5. As recently as 1965, Carleton Coon assumed the most fundamental division in humanity was along the Movius line, with East Asians off by themselves and whites and blacks more closely related to each genetically than to East Asians.

    It’s interesting how population genetics has moved the orthodoxy in a less politically correct direction toward the fundamental division being sub-Saharan Africans vs. Eurasians.

  6. “What isn’t easily explained is why sites after 50,000 years ago should be susceptible to this same path-dependence — particularly since they should derive some or most of their ancestry from Africa/West Asia.”
    “Personally, I keep coming back to ecology.”

    It’s been said that the lack of raw materials and low quality of the material was a restraining factor on Levallois, but I guess the only progress in this field is actually dependent on findings of another field, genetics, which can at least in theory let us know which way(s) modern and prehistoric populations arrived into East Asia, either via Central Asia which seems to have Levallois or via the southern coastal route which seems to have forgotten it.

    “particularly since they should derive some or most of their ancestry from Africa/West Asia.”
    Only some? I agree that shoveled incisors are perhaps a trait “Mongoloids” got from erectus, but the erectus component overall should be very low based on what we know from genetics. Not one phylogenetically-distant sample of mtDNA and Y-Chromosome have been discovered yet. All fall into a sub-set of East African/Eurasian lineages.
    My theory on how the above seemingly contradictory situation can happen is this:

    1. The erectus admixture must’ve occurred in a very small population.
    2. The uniparental markers died out in this bottleneck but the autosomal genes remain.
    3. The shovelling trait was selected in severe ecological conditions.
    4. This “Mongoloid” population at some point started to expand exponentially.

  7. I think the same Movioid pattern can be established for the populations of Homo sapiens sapiens. We see “delays” in the appearance of aspects of the modern human behavioral package in Australasia and especially in the Americas. Clovis points in North America are so much like Solutrean points in Europe, only 10K years younger. The very absence of ancient archaeological signatures in the Americas may not mean the absence of humans earlier than 13.5K years but rather low population densities and technological stagnation for a long period of time. Low effective population size in Asia and the Americas, as compared to Europe and Africa, is expressed in the reduced levels of haplotype and other measurements of diversity. At the same time, the geographic range of haplotypes is broader east of the Movius line (mtDNA haplotypes C and D, for instance, are found from India and Scandinavia to Tierra del Fuego, while Y-DNA C haplogroup is found from Australia to North America) suggesting long-term stability and lack of mutational innovations. At the same time, the levels of LINGUISTIC diversity are much higher east of the Movius line than in Africa or Europe, with America and Papua New Guinea language families encompassing 3/4 of world linguistic diversity. This again suggests great antiquity and low population size east of the Movius line, as languistic diversity shows no population-size dependency.

  8. @ John Hawks: I’m not wholly convinced by the ecological reason. Primarily because East Asia is quite large and covers a variety of ecological niches; it’s not a uniform area. I suppose, really, the argument here is whether or not the ecological conditions trumps the demographic underpinnings. Also, it may not be one factor over another that explains why the patterns of handaxes east of the Movius line tend to be geographically sparse; instead, it may very well be the combination of partly ecological factors and partly the demographic limitations surrounding cultural and technological development. I think there are three important factors to consider in this particular context:

    1) Was there available access to significant quantities of high-quality, fine-grained stone for tool production in these areas east of the Movius Line? Obviously, blade technology is quite dependent on fine-grained stone such as flint, chert, obsidian etc. Furthermore, did hominin populations rely on locally-sourced materials, such as wood or bamboo, for tool manufacture.

    2) What are the specific functions for this stone tool technology? For instance, if the initial colonisation followed a costal route, then these technologies may have needed to adapt for the exploitation of coastal resources.

    3) Lastly, there are the issues of demography and the potential role of serial founder effects as relatively small population units migrated eastward. Do we see instances where serial founder effects not only operate on the genetic features of populations, but in addition leads to a loss in the complexity and diversity of cultural and technological patterns as the result of an increasing distance from the demographic point of origin?

    As for the point about those Flores hominids: I don’t know enough about them to really make an informed comment. For the moment though, I’d say that having a small population is not the only factor. If they had a sufficient level of density, then this may have led to stronger and more regular instances of social interconnectedness. So, as the probability of encountering increases, so does the ability to maintain certain forms of technology.

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